To misinterpret fear as surprise... That must be one bad surprise.
"So sounds like if the game is a global audience, the amount of morphs or bones around the eye need to increase as well as having animated pupil dilation. "
I don't think there needs to be a big increase in eye animation. The article states that the emotions are misinterpreted because the movements of the eyes are ambiguous between certain expressions. So maintaining realism, other cultures just have to deal. There's also the fact that likely there is voice over which is also conveying a lot of emotion, so coupled with the facial expression the message should be understood.
Interesting article though, good read.